An alpaca at the centre of a legal row with the government has been killed.
Geronimo twice tested positive for bovine tuberculosis, and the Department of Food, Environment and Rural Affairs (Defra) ordered him to be euthanised to stop the spread of the disease.
Owner Helen Macdonald disputed the results and mounted several High Court challenges to save him.
"I am absolutely disgusted by this government. These are barbaric actions," she said.
Defra staff dressed in blue overalls, goggles and masks, accompanied by police, arrived at her farm in Wickwar, near Bristol, at about 11:00 BST and led Geronimo away, with the scene broadcast on an online live feed of the enclosure.
A spokesman confirmed the destruction warrant had been carried out after Geronimo was taken away from his pen and loaded into a trailer.
The vehicle then left the scene under a police escort, with a number of protesters watching.
Ms Macdonald said the government had refused to act in good faith and had ignored her pleas for constructive dialogue.
"We know now they have been stringing us along for the last week, fobbing us off by saying people are on holiday and would get back to us this week," she said.
"In fact, all the time they were simply planning to murder Geronimo. This is yet another appalling demonstration of bad faith and duplicity by the secretary of state and everyone at Defra.
"Geronimo has been manhandled out of my farm. He was rounded-up and dragged into a horsebox."
More than 140,000 people from around the world had signed a petition addressed to Prime Minister Boris Johnson to stop Geronimo's destruction.
Wildlife television presenter Chris Packham and the prime minister's father Stanley Johnson also gave their support.
A Downing Street spokesman said: "It's obviously highly distressing for someone to lose animals to TB and that's a situation that farmers sadly have to face.
"Our sympathies are with Ms Macdonald and any others that are affected by this terrible disease."
Ms Macdonald insisted Geronimo was a "perfectly healthy animal", adding: "Boris apparently is sympathetic, well I don't need his sympathy.
"I needed him to stop this in its tracks when he had the opportunity and look what's happened."
Chief veterinary officer Christine Middlemiss said it was a "terribly sad situation" but they had to follow the scientific advice and cull animals that tested positive for bTB in order to minimise its spread.
"Not only is this essential to protect the livelihoods of our farming industry and rural communities, but it is also necessary to avoid more TB cases in humans.
"No-one wants to have to cull infected animals if it can be avoided," she said.
A Defra spokesman said a post-mortem examination and a study of tissue samples would be carried out by veterinary pathologists from the Animal and Plant Health Agency and the process could take up to three months.
Ms Macdonald has called for Defra to allow for an independent witness to attend the examination alongside the pathologists.
'Devastation'
Defra said 28,356 cattle were compulsorily slaughtered in England in the 12 months to March 2021, after testing positive for bovine tuberculosis.
During 2020, a total of 205 camelids - animals such as camels, llamas and alpacas - were culled.
Defra said: "Bovine TB remains one of the greatest animal health threats to the UK, causing devastation for hard-working farmers and rural communities.
"We are pursuing a range of interventions in cattle and in badgers to eradicate the disease by 2038.
"If we do not take action, TB could pose an increased risk to animal and human health and to the international image and reputation of our cattle industry."
Geronimo timeline
- Geronimo is brought to England from New Zealand in August 2017 by his owner Helen Macdonald
- The alpaca twice tests positive for bovine tuberculosis in August and November that year
- He is put into isolation away from the rest of the herd at the farm in Wickwar
- The government applies for a court order in July 2018 to have him destroyed
- Geronimo is given a stay of execution, with a deadline of the end of August for his slaughter
- Ms Macdonald seeks a judicial review claiming new evidence shows the animal is healthy, marking the start of a series of lengthy legal battles
- In November, Ms Macdonald wins the right to a review at the High Court in London
- A hearing gets under way in March 2019 and Ms Macdonald claims government experts relied on "flawed science"
- The case is dismissed in July 2019
- Ms Macdonald starts an appeal and an order is made preventing Geronimo's death pending the application
- She takes out an emergency injunction to delay a warrant to cull the animal before an appeal hearing on 29 July 2021
- The case is once again dismissed. A judge agrees to delay the start of a second execution warrant until 5 August, which is later extended to 4 September
- Geronimo is euthanised on 31 August
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